South Alabama’s second football scrimmage of the spring featured plenty of highlights in the running game.

The Jaguars’ first-, second- and third-team offenses piled up 388 yards* and four touchdowns rushing, part of a 714-yard, 28-first down day by the offense. South Alabama had several explosive plays on the ground, breaking off runs of 95, 40, 36, 30, 26 and 23 yards during a scrimmage that totaled approximately 112 plays at Hancock Whitney Stadium.

(* — stats are unofficial, compiled by AL.com)

“We were making some plays,” South Alabama coach Major Applewhite said. “That’s linemen blocking, that’s backs breaking tackles, that’s wide receivers blocking. We’ve got to go back and look at (the scrimmage film), see if some of those are missed tackles, but pleased with the way (all the backs) ran.

“ … There was an emphasis when we were moving the ball in certain parts of the scrimmage where we were trying to push the tempo and go fast, then there was other parts of scrimmage where we were trying to give a lot of exotic formations and motions, and I thought we had a good scrimmage in terms of the situations, the effort. I’ve got to go back and really watch it to see how physical we were, see what our finish was, but it felt like guys were competing.”

Junior-college transfer Keenan Phillips ran for 121 yards on just three carries, including a 95-yarder on which he was caught at the 5-yard line by cornerback Ricky Fletcher. PJ Martin added 96 yards and a 36-yard touchdown during a 2-minute drill on seven carries while splitting time with the first- and second-teams, and also had a 40-yard run against the first-team defense.

Starting running back Kentrel Bullock ran for a 30-yard touchdown, while Iverson Celestine scored from 12 yards out against the second-team defense and third-teamer Chris Holifield on an 11-yard run during red-zone work. Bullock’s touchdown was a bruising effort, on which he broke at least two tackles inside the 5-yard line.

“It feels good, I can’t lie,” center Malachi Preciado said of all the explosive runs. “Just to think, if you guys were here last spring, we weren’t getting a lot of those. The steps we’ve taken to see our running backs really go and unload, it feels good.”

The South Alabama passing game was also effective, with four quarterbacks combining to complete 37 of 59 attempts for 326 yards and two touchdowns with no interceptions. Starter Gio Lopez was good on 18 of 26 passes for 155 yards, including touchdowns of 38 and 11 yards to Devin Voisin, who had 65 yards on five receptions overall.

Second-teamer Bishop Davenport completed 13 of 19 passes for 134 yards, while third-team QB Jared Hollins was 5-for-12 for 28 yards. Walk-on Bubba Thompson played one series, completing one of two passes for nine yards, but also breaking off a 12-yard run.

Anthony Eager added six receptions for 42 yards, while Bullock and Trent Thomas had four catches each. Jaguars kickers made both of their field goal attempts during red-zone work, Davis Little from 30 yards away, Hamilton DiBoyan from 33 yards out.

The South Alabama defense had a few highlights, including Fletcher’s touchdown-saving tackle, a pass breakup by Wesley Miller and a sack and tackle for loss by Tyler Carter. The Jaguars totaled two sacks, two pass breakups and three tackles for loss, but for the second straight week did not force a turnover.

South Alabama’s defense did go 2-for-4 in the red zone, forcing two field goals. The Jaguars also got off the field on one of two two-minute drives.

“I saw a lot of chemistry out there,” linebacker Darius McKenzie said. “A lot of guys flying through the ball, swarming, and just great energy by defense, offense. Just a great day overall, seeing everyone fly to the ball.”

South Alabama returns to practice on Tuesday morning, then will work Thursday night leading into the Spring Showcase next Saturday at Hancock Whitney Stadium. The Spring Showcase — which begins at 10 a.m. and takes the place of a tradition spring game — features an hour-long open practice and skills competition, along with an Easter Egg hunt, the Jaguars’ annual surplus equipment “yard sale” as well as a post-practice autograph session and kids’ clinic.